In Lejeuneaceae vegetative branches normally originate from the basiscopic basal portion of a lateral segment half, as in the Radulaceae, and the associated leaves, therefore, are quite unmodified.
The Jubulaceae have a leaf whose lobule, usually transformed into a water-sac, is normally very narrowly attached to the stem and to the dorsal lobe; indeed some Frullania taxa reproduce vegetatively by dropping the dorsal lobes, but not the lobules, and Neohattoria has caducous lobules but persistent lobes.